The segmentation unit performs two operations simultaneously: adding the segment base to produce the linear address and comparing the effective address against the segment limit. Both use dedicated 32-bit adder/subtractor circuits.
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He says the results could change predicted extinction dates for Emperor penguins.
Real game engines use this pattern (or its 3D cousin, the Octree) for broad-phase collision detection: the quadtree quickly identifies candidate pairs, and a more expensive narrow-phase check tests the actual geometry.